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Messages - SoCal2warm

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126








Pomegranate with rare Passe Crassane Pear (actually a quince-pear hybrid) in the background



That's a Pisang Ceylon you see in the background, a less common but very worthy variety



A Yoshino sakura can grow in climate zone 10, apparently, but only if grown own-root
Some things you don't find out unless you try



May not look like much but this is actually Western Maidenhair Fern. These things are notoriously hard to transplant. It's said that the Western variety (as opposed to the Northern Maidenhair which is nearly identical) is a bit better adapted at being able to handle the dry summers of the West coast, even though both these Maidenhairs are very vulnerable to drying out and grow best near waterfalls, or sloped hills that only get morning sun. Beautiful shape, and the Western variety is virtually impossible to find in nurseries outside the Pacific Northwest.





Here's the lychee and Karp's Sweet again



Oregold



Pomelo and Sweetheart lychee



Some fragrant flowers, Madagascar Jasmine



127
Heinrich and Ilya11, you are both being illogical and drawing faulty conclusions.

Heinrich, you said "Nucellar seedlings are never derived from meiosis". I did not imply that they were.
In fact by the very fact that they were (presumably) not derived from meiosis, you can conclude something about meiosis.
If there were no triploids resulting from meiosis, what does that tell us? And it would certainly seem to indicate something about the origins of that tetraploid they found.

Ilya11, go back and read over everything I posted. I never said that triploid plants gave rise to haploid offspring that were alike. You seem to be assuming I said something that I did not.

Ilya11 has still failed to give any reasoning why what I wrote is wrong.

Pollination of Oroblanco with diploid clementine pollen is giving only diploid plants due to unfavorable completion of diploid gametes   with haploid ones.
(Citrus Genetics, Breeding and Biotechnology,edited by Iqrar Ahmad Khan, p.205)
That's not necessarily true. That's just an assumption they made. 

My alternative theory offers an explanation. If Oroblanco is not producing any diploid gametes then of course there would not be triploid offspring. No haploids are resulting because the Clementine pollen is not triploid and is therefore unable to induce gynogenesis in the Oroblanco, and besides, the female parent being triploid might have something to do with it (apparently it may take triploid pollen pollinating a diploid plant for this effect to happen). Sometimes, in certain species, nucellar seed formation can be induced by pollination, when the resulting embryo is not very strong as a result of chromosomal differences/abnormalities, so if the majority of the pollen is not triploid, that could be an explanation for the lack of nucellar formation.

128
Nucellar seedlings are never derived from meiosis. Therefore, you canīt conclude from nucellar seedlings anything about meiosis.
I meant that, because all the triploids identified were nucellar, that seems to indicate no triploids resulted from meiosis, and by implication it seems meiosis of a triploid set is unable to produce a diploid gamete (or at least it's very rare). So if there were no triploids resulting from meiosis but they identified one tetraploid, it's logical to conclude that that tetraploid was very unlikely to have been a result of triploid meiosis.

For that to happen, there would have had to have been two different triploid sets that each split into an intact diploid, and then those diploid gametes would have had to have fused together to produce the final tetraploid. Not a very high probability of happening, especially since presumably no triploids formed from diploid gametes.

129
Hey guys, I know this post is super old, but I was wondering how does Luc's Garcinia taste in comparison to an achacha?
Probably a little bit better

And how would you compare it to a purple mangosteen?
Depends on personal preference.

Here's the best analogy I can think of. For lychees, do you prefer Brewster or Mauritius? If you prefer Mauritius I believe there's a higher chance you will prefer achacha/Luc's over purple mangosteen.

130
Citrus Buy, Sell, & Trade / Re: Wanted: yuzu
« on: August 20, 2017, 03:13:53 AM »
La Crescenta Nursery carries yuzu, although they're a bit pricey. Only 5 hours away from you.

131
Cold Hardy Citrus / Re: breeding cold hardy pomelo
« on: August 19, 2017, 04:23:21 PM »
Just wanted to add: it's not really true that pomelos are always that cold vulnerable. Limes are probably more cold vulnerable than pomelo trees. Pomelo might fall somewhere close to lemon. To some degree it depends on the cultivar variety. Pomelos usually come from tropical countries like Thailand, Malaysia, Vietnam (particularly the southern half), although there is a variety in Thailand that grows in a slightly more mountainous area in zone 9b and is known to be just a little more cold hardy. Then there are the "Chinese grapefruits" which I personally suspect might actually not have 100 percent pomelo ancestry, but who knows... They are grown in Southern China. The winters there can be cold, if not freezing, going all the way into Spring, cold and wet. (Though after that it's hot and humid, great growing conditions) It is zone 10, but the early Spring is far colder and wetter than what those in zone 10 California experience. Each climate zone has its own subtleties I suppose.

The pomelos towards the more cold hardy end of the spectrum probably have as much cold tolerance as the average orange, if I had to guess, while there's certainly many pomelos in Thailand that wouldn't be happy with anything lower than zone 10. (necessary heat for ripening is a different matter)

I suspect most pomelos in Thailand and Malaysia would be happier in zone 11 than they would in zone 10. Or to be a little more exact, happier in 10b bordering on the edge of 11a than they would be growing right on border between 10b and 10a. Chandler is probably just a tiny bit more cold hardy than the average pomelo variety in Asia (but I could be wrong about this).

Going back to the Chinese grapefruits really quick, they are probably all or mostly pomelo, and are classified under Citrus maxima. I'm just thinking there might be somewhere like 10 percent C. reticula in there, since they are sweeter and do have a thinner rind than other pomelos. "Chinese grapefruit" really isn't the best terminology, that's just the common English translation, from a time Americans were much more familiar with grapefruit than having any idea what a pomelo was. In Asia though they're both considered the same thing ( "Chinese grapefruit" and pomelo I mean), they don't think of the different varieties as a fundamentally different fruit. Maybe I should have best avoided using the word "grapefruit" altogether to avoid confusion and just called it Chinese-type pomelo. (by using the word grapefruit I was just trying to emphasize that it might not be completely C. maxima)

132
Cold Hardy Citrus / Re: breeding cold hardy pomelo
« on: August 19, 2017, 01:56:20 PM »
Here's a Yuzu tree in California



found it after scourering several local nurseries in the area

133
An interesting paper:
https://ejournal.sinica.edu.tw/bbas/content/2009/3/Bot503-06.pdf
Thank you so much.

If I can summarize the results here: They looked at the seeds of triploid citrus (this particular cultivar having resulted from a tetraploid cross between Hamlin orange x rough lemon, which then was crossed with diploid tangerine) which was open-pollinated, and out of 25 well-developed seeds, 13 were triploid, 10 were diploid, 1 was tetraploid, and 1 was aneuploid. What was particularly interesting was that all of the triploids appeared to be nucellar. This suggests that the likelihood of triploid meiosis being able to produce a complete haploid gamete is far higher than a diploid gamete.
Assuming this is the case, then presumably the single tetraploid seed must have resulted from an unreduced triploid gamete being sexually combined with a normal haploid gamete.
It seems then that the probability of a complete whole diploid chromosome set being able to escape from triploid meiosis is very unfavorable, all the triploids resulting from a triploid fruit will be clones of the parent. (The triploids will be the offspring with very few seeds)

134
Temperate Fruit Discussion / Re: Apple trees
« on: August 15, 2017, 11:43:27 PM »
There may be other possible reasons, like if one tree is very young, or if one tree gets a lot more shade than the other tree.

135
Disclaimer is right, it is wrong.
Triploids never give haploid plants ; theoretically  their gametes  are only  haploid, diploid and triploid.
I believe you were misunderstanding. Triploid pollen can induce the formation of haploid embryo from the female fruit parent; the single chromosome set would always come from the female parent in such a case.

Quote
This study reports haploid plantlet regeneration through gynogenesis in Citrus clementina Hort. ex Tan., cv. Nules, induced by in vitro pollination with pollen grains of Oroblanco, a triploid cultivar of grapefruit. It indicates that parthenogenesis induced in vitro by triploid pollen can be an alternative method to obtain haploids in monoembryonic cultivars of Citrus.
https://link.springer.com/article/10.1023/A:1010627310808


There are a few possible interpretations to these findings (not discussing the link above now). Unfavorable competition of one pollen type was only listed as a possibility, as well as a possibility all the triploid seeds (when they form in Oroblanco fruits) didn't result from pollination.

136
So a quick summary, here is what I gathered from the information
(disclaimer: this could be completely wrong)

When Oroblanco pollinates a normal diploid, you get a haploid (at least the vast majority of the time), or in other circumstances apparently, maybe depending on the cultivar, you will get all diploid
When Oroblanco self-pollinates, you get half diploid and half triploids (and doesn't seem to cause gynogenesis probably because the ploidy of the fruit parent is the same as the surrounding pollen tissue)
There is probably a tiny ratio of tetraploid but those seeds are very underdeveloped and will not germinate on their own.
When a normal diploid pollinates Oroblanco, you probably get diploid and triploid

137
After several hours of research I came across something.
P. Ollitroult found half diploid and half triploid among the seeds of 'Oroblanco'.
Citrus genetics, breeding and biotechnology, Iqrar A. Khan, p205

Only diploids were found in the progeny of clementine fertilized by Oroblanco pollen, the same source says. But I wonder if they checked to see if the next generation were even hybrids, since very likely the triploid Oroblanco pollen might just have been inducing parthenocarpy. Edit: oops, if that was the case, one would expect haploid offspring (see gynogenesis). Another potential explanation, I did read in some citrus cultivars (particularly many oranges) the surrounding nucellar tissue in the embryo is highly sensitive to triploidy and will abort.

Also, while it is true triploid seeds are most often found in shrivelled up smaller seed (which also causes low viability), this is not necessarily true when the fruit parent happens to also be a triploid. The poor seed development is caused by differences in ploidy between the developing megagametophyte and surrounding nucellar tissue.

138
Some of you may be aware that Oroblanco and Melogold grapefruits are triploid, meaning they have 3 sets of chromosomes rather than the normal 2.
So a natural question that arises is what would happen if you tried to breed a triploid citrus with something else? What would you get.
Although Oroblanco is normally considered seedless, I found several fully formed large seeds inside one the fruits I just bought. I'm assuming all/most of these seeds are nucellar (clones of the parent) but it got me thinking.

I've been doing a lot of research, and wanted to present to you what I've found so far, in an attempt to answer the question. I'm no expert in this area.

Basically, when a triploid like Oroblanco pollinates another citrus variety (generally a monoembryonic variety) it induces the formation of seed. These seeds will be haploid, having only a single set of chromosomes from the female fruit parent. Most of the gametes produced from a triploid will have unevenly split chromosomes during meiosis, but a smaller portion will also be diploid and haploid. That means that pollen from a triploid parent could contribute either one or two sets of chromosomes. Assuming that the female parent is diploid, this will result in the seed either having 2 or 3 sets of chromosomes. So you could get a normal zygotic seedling, or you could get another triploid. The triploids will have a lower probability of having fully formed seeds, so the majority will not survive. (Embryo rescue techiques are usually employed to be able to grow these underdeveloped seeds, but in a small number of cases the seeds will be well formed enough to be able to germinate on their own)

It is also possible for a triploid parent to produce an unreduced gamete but this comparatively much more rare, so in that case the gamete would be able to contribute 3 sets of chromosomes.

Haploid citrus plants are normally feeble and slower growing, generally sterile, and right now I cannot find any references to a haploid citrus plant ever having produced fruit. But for breeding purposes, they can be reconstituted back to normal diploids by treating the growth on bud shoots with colchicine, and then later taking a graft from that offshoot.

This still leaves so many questions but it's a start to understanding.

Quote
Triploid plants can and do go through meiosis (cite). Of course, their fertility is lowered with most gametes being aneuploids (uneven number of chromosomes), however, triploid plants also produce small numbers of euploids (1x, 2x, 3x)(Fig 1 of the above cited source).

This should make you wonder why we don't often observe seeds in triploid fruit since they can make viable gametes (1x, 2x, 3x). This seems to be purely driven by the number of female gametes (ovules) produced vs. the number of male gametes (pollen grains) produced. Plants produce many more pollen grains than ovules making it much more likely that a euploid pollen grain will be produced. Contrasted by the small number of ovules produce making it very unlikely that a euploid ovule will be produced.

The fertility of triploid plants is the foundation for one of the most used mechanisms for polyploid formation called the triploid bridge. The triploid plant produces 1x, 2x, and 3x gametes that combine with the 1x gametes produced by a diploid plant to form 2x, 3x, and 4x offspring
 

I could well be wrong about most of the seeds in Oroblanco being nucellar. That would be interesting.

139
Citrus General Discussion / Re: The History Of Citrus Fruit
« on: August 12, 2017, 01:38:09 PM »
I have no idea what the cone of an umbrella pine (Pinus pinea) looks like, but I know the tree grows all around the city of Rome.
Apparently it bears a resemblance to pineapple when you try to depict it in a mosaic.

140
Citrus General Discussion / Re: whole new subcategories of citrus
« on: August 12, 2017, 01:14:06 PM »
I did not say any of those fruits you just listed have papeda ancestry.

The problem is, while the common citrus have been extensively sequenced, the more obscure ones have not. This can result in misleading correlations in the breakdown. It's more likely to show a citrus that has more recorded markers (say citron for example) than another uncommon citrus (say papeda) with only a few recorded markers. You can't really have a good idea whether a hybrid has some type of papeda ancestry unless you have looked closely at the DNA of that papeda and are considering a lot of markers from that species in the comparison.

Now of course most of the common citrus varieties don't have any papeda ancestry, if that's what you thought was misleading. I'm sure those tests are very accurate if you look at the citrus being sold in a supermarket, or unusual traditional cultivars in Europe, but if you're analyzing unusual citrus in Asia it can be a different story.

141
Citrus General Discussion / Re: The History Of Citrus Fruit
« on: August 12, 2017, 12:06:36 AM »
Just a an aside here, does anyone know what the other fruits in this fruit basket could be?
I would say at first glance: figs, quince, grapes, pomegranate and pineapple.
But, wait a minute, pineapple? In the first century BC in Rome?
What is going on here?
Most scholars believe the mosaic depicts the cone of an umbrella pine (stone pine), from which edible pine nuts come from. They were very popular in Italy.

142
Citrus General Discussion / Re: The History Of Citrus Fruit
« on: August 11, 2017, 10:55:35 PM »
East Asia has an entirely different history of citrus than Europe, and by extension the United States, does. Citrus travelled in different directions, through different climate areas, and so different types were preferred in different climate conditions. Citrus was slow to disseminate across vast distances, but citron had already reached the Mediterranean in ancient times, possibly through the Persian Empire. The orange was next, probably because the fruit had a longer shelf life to be carried by travelers, but had more cold tolerance than pummelo. Initially it was the sour orange, but the sweet orange came very soon thereafter. Although sour orange was known by the Romans, sweet orange didn't really become widely known in Europe until the Fifteenth Century, and orange growing was concentrated in Italy and Spain. Already in the Tenth Century muslim rulers in Spain had planted sour orange trees in their gardens, and by the Thirteenth Century there were orange groves in Spain.

The Northern part of China has always preferred mandarins, hence the name, while in Southern China they have long had "Chinese grapefruit", which is all or mostly pummelo, probably originally brought from Vietnam. In Japan, mandarins (and what are the equivalent of tangors) originally came from Northern China (this probably took place mostly during the Tang dynasty), while Chinese grapefruit came later (less than 400 years ago), from a Cantonese ship from Taiwan.

Pummelo was probably endogenous to the area around Malaysia and the southern regions of Thailand and Vietnam. Mandarins probably originated somewhere in Southern China.

In China, the citron is called "fragrant ball" (this is also the same name which the Ichang lemon is known by in China, although the two are not so much related). The fruit known as Yuzu in Japan is known as "fragrant orange" in China.

143
Citrus General Discussion / Re: The History Of Citrus Fruit
« on: August 11, 2017, 10:18:33 PM »
The pomegranate juice is fantastic!  But yes, I'm always disappointed when I buy one and after all that work realize a bottle of Pom juice would be far easier.
A bottle of Pom juice from the supermarket doesn't taste anywhere near as good as real fresh squeezed juice. Pomegranates are really one of the fruits you have to squeeze fresh to be able to enjoy it, the flavor quality apparently deteriorates a lot with longer term storage and processing. And there's a certain way you have to make the juice too: first separate out the arils from the rind and then press those arils for the juice, because both the seeds inside the arils and the inner white rind can impart off-flavors into the juice if it is made by other methods.

144
Citrus General Discussion / Re: whole new subcategories of citrus
« on: August 11, 2017, 07:00:54 PM »
Those DNA studies are pretty good at telling how much C. maxima relative to C. reticula a citrus has in its ancestry, but when it comes to differentiating between citron and different papedas, the standard DNA tests are not that good. Apparently the many of the markers being looked at in the analysis are not unique to just one of these more obscure citrus species, so when you see those broadbased diagrams across numerous citrus varieties, you have to take them with a grain of salt. (Unless someone ran a DNA analysis specially focusing on one of these papedas and their immediate hybrids)
You can tell this because many times those analysis results will be intuitively wrong in specific instances. What might really be a yuzu ancestor will show up as directly Ichang papeda. What might really be a papeda ancestor might show up in the analysis as likely citron. It's not the tests themself that are flawed, but rather the poor selection of genetic markers to be able to distinguish some of the less common citrus species.

145
Citrus General Discussion / Re: whole new subcategories of citrus
« on: August 11, 2017, 01:18:12 PM »
Khasi papeda looks incredibly similar to kaffir lime. I don't think kaffir lime is a pure species though, probably a hybrid between sour mandarin and papeda, or something along those lines. Or quite possibly a hybrid between khasi papeda and citron.
Lemon is a hybrid between citron and sour orange (bitter orange). Meyer lemon is hybrid between citron and mandarin, or something very close to mandarin, much closer to mandarin than orange. Ponderosa lemon I have read several places is supposedly a hybrid between lemon and citron, but I think it more likely it originated from a lemon pollinated from grapefruit or shaddock.
I'm not sure what regular limes are though. Supposedly a hybrid between C. micrantha and citron. There are different types of limes so some might have additional ancestry.

There's a lot of information here and I'm typing this off very quick, so some of the information might not be 100 percent completely literally accurate.

146
Citrus General Discussion / Re: Favorite citrus options...
« on: August 10, 2017, 11:04:41 PM »
Well if you're looking for some unusuals I might recommend a wekiwa tangelolo or dekopon. Satsuma mandarin is always a good one to have in the yard or pummelo (if you like those). Oroblanco is popular with home growers because they are sweet and juicy.

It's hard to tell you what to get because in many cases it can be a matter of personal preference.

147
Citrus General Discussion / Re: Grapefruit Oil
« on: August 10, 2017, 02:39:44 PM »
Pomelo oil is similar to grapefruit but has a cleaner more floral smell, without the slightly sewer-like aspect of grapefruit. Unfortunately pomelo is substantially more commercially expensive than grapefruit because there's a much more limited geographic extent in the U.S. where it can be grown.

148
Citrus General Discussion / Re: whole new subcategories of citrus
« on: August 10, 2017, 06:15:16 AM »
I still think micrantha is just a form of hystrix but didn't know it is a progenitor of modern limes. I also expected maxima to feature more prominently and medica to feature less.
I like the way you think, but I don't think that's true here. I seem to remember reading that kaffir lime was a hybrid of some other type of papeda (I can't remember what it is right now) from which it derived its cold-hardiness. (It was some weird one, not as cold-hardy as Ichang papeda, but maybe I'm not remembering any of this right)
I think kaffir limes are descended from a different papeda ancestor than common limes. Isn't kaffir lime about as cold-hardy as the more cold-hardy oranges? While common limes are the most sensitive out of all the different citrus groups to cold, slightly more sensitive than common lemons.

Edit: I think it may have been C. latipes, also known as Khasi papeda.

But yes, it's well known that limes are descended from C. micrantha. Another little uncommon fact, C. micrantha is incredibly high in furanocoumarin levels, over four times higher than bergamot! Probably not a good idea to put the essential oil on your skin.

So lemon's primary ancestor is citron, for lime it is C. micrantha, and for kaffir lime it is Khasi papeda (assuming I'm remembering right)

I do suspect citron, C, micrantha, khasi papeda (and Ichang papeda too) are closer related to each other than they are to C. reticula or C. maxima, but that's just judging by morphological appearances.

149
Citrus General Discussion / Re: breeding - what can I do with citron
« on: August 09, 2017, 10:50:10 PM »
Thank you.

The seeds are already sprouting, and have tiny little leaflets. I have two different types of Etrog citron: an unnamed one that I assume is the most common, and Yemeni Giant citron, the latter of which can often get bigger than a pomelo, from pictures I've seen (although it doesn't have any pulp inside).

There are actually many different types of citron, I was surprised when I started doing research.
In China it is known as Xiang Yuan. This is actually the same word as Shangyuan (just a different romanized spelling), which just translates as "fragrant ball". Shangyuan in English circles is most commonly associated with the Chinese citrus known as Ichang Lemon (not to be confused with Ichang papeda). And Chinese citron is not Ichang lemon (I don't think Ichang lemon even has any citron ancestry, unlike the European lemon). It can all be rather confusing.
The Chinese citron is commonly used to make a type of jelly tea. (in fact I have a Chinese friend who offered me some from his refrigerator, now that I remember)

These language misunderstandings can be confusing. When the Japanese first encountered Yuzu during the Tang dynasty, they named it after the Chinese word for pomelo.

Well that's your little world history cultural lesson for the day.

150
Tropical Fruit Discussion / Re: Lychee problem - please help
« on: August 08, 2017, 01:47:48 PM »
I have had a hard time with lychees. I thought the brown burnt leaves are from salt either in the water
or fertilizer. I use to use synthetic fertilizer and on the label it says not less then 7% chlorine.  I finally got
a mauritas  lychee doing well and 3 years ago we had a 30F night and it killed it. Currently I have a ohia and
it is doing well. Last fall the leaves all burnt and I attributed it to too much fish emulsion? The only other thing
could have been a reduction of water? It has flushed a couple good growths and most of the burnt leaves
have shed. I would water daily and if possible use rain water. Check your previous fertilizer for the contents and now
I use manure and only organics like coffee grounds and compost. When you go to fruitscapes the lychees look so
damn good in the pots. They use a slow release fertilizer and water daily. One of these days I will crack the lychee
code.
That's not an uncommon problem with lychee. The problem where you are could be sandy soil. Make sure the soil is rich and loamy and holds water well. Also, you may want to think about using shade cloth, or planting another fast growing plant to provide shade to the lychee tree, at least for the first couple of years. It takes time for the root system to become established. Until then, the tree can be vulnerable to drying out. It may take consistent watering to keep the soil moist. Salt may also be an issue, like you said. Sometimes the soil in Florida can have a lot of salt in it. That may mean you will have to dig a very big hole and replace the soil with bagged soil and compost. It's also possible you may be giving it too much fertilizer, that can also cause leaf burn.

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