Acidless citrus varieties do initially synthesize citric acid during early fruit development. In certain acidless cultivars such as Sucreña, this acid is subsequently broken down through elevated activity of aconitase enzymes, as demonstrated by increased expression of CcAco1 and CcAco2 (Terol et al., 2010).
Therefore, there is no net energy saving associated with the acidless phenotype. On the contrary, there may be an additional metabolic cost due to the energy required for both the synthesis and later degradation of citric acid.
Terol, J., Naranjo, M. A., Ollitrault, P., & Talon, M. (2010). The aconitate hydratase family from Citrus. BMC Plant Biology, 10, 222.
https://doi.org/10.1186/1471-2229-10-222