I read somewhere that these incompatibilities depend on the particular clones of lemon varieties due to the presence of virus infections in them.
The UC germplasm is supposedly virus free, but I guess I don't know if that was the case when they produced that chart. I would love to read a study that discussed that issue though, if you can track it down.
There are many other citrus incompatibilities though. E.g., grapefruit/C-35. Are you suggesting that every single instance of incompatibility in citrus is due to virus issues?